The book opens with the discovery that Andromeda, the enemy galaxy of the first novel, has discovered the secret of involuntary hosting: a sufficiently higher-Kirlian aura can in effect possess an individual of a lower-Kirlian aura. This has enabled Andromeda to secretly infiltrate the highest levels of government in Sphere Sol and its allies and resurrect its plot to steal the energy of the Milky Way.
Melody of Mintaka, a direct descendant of Flint of Outworld and his Andromedan nemesis, has a Kirlian aura of well over 200. She is pressed into service to "possess" and interrogate a captured Andromedan transferee. Melody, hosted in the young and beautiful body of Yael of Dragon, must like her ancestor Flint find a way to defeat the Andromedan threat and save the galaxy.
The mysterious Ancients are present again in the form of their artifacts and sites. The themes of Tarot and of various myths of Sphere Sol (in this case that of Perseus and Andromeda) are carried throughout this novel as well -- for example, the interSphere fleet of starships has forms analogous to the Tarot suits of Disks, Cups, Wands and Swords.